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Dr. Gareth Griffith



Davey, H.M., Cross, E.J.C.,  Kell, Oliver etc. Griffith, G.W. (2012). Genome-Wide Analysis of Longevity in Nutrient-Deprived Saccharomyces cerevisiae Reveals Importance of Recycling in Maintaining Cell Viability. Environmental Microbiology, In press.

Bussell, J., D. Gwynn-Jones, G.W. Griffith and J. Scullion (2012). Above- and below-ground responses of Calamagrostis purpurea to UV-B radiation and elevated CO2 under phosphorous limitation." Physiologia Plantarum, In press.

Griffith, GW (2012).  Do we need a Global Strategy for Microbial Conservation? Trends in Evolution and Ecology, 27(1),  pp. 1-2.  DOI: 10.1016/j.tree.2011.10.002 (published online 8th Nov 2011). Download the pdf.

Griffith, G.W., Roderick, K., Graham, A., Causton, D.R. (In press). Sward management influences fruiting of grassland basidiomycete fungi. Biological Conservation, 145, pp. 234–240. Download the pdf.


Kumar S, S.S. Dagar, A.K. Mohanty, S.K. Sirohi, M. Puniya, R.C. Kuhad, K.P.S. Sangu, G.W. Griffith & A.K.Puniya, (2011). Enumeration of methanogens with a focus on fluorescence in situ hybridization. Naturwissenschaften DOI 10.1007/s00114-011-0791-2. Download the pdf.

Griffith, G.W., Baker, S., Fliegerova, K., Liggenstoffer, A, van der Giezen, M., Voigt, K. and Beakes, G. (2010). Anaerobic fungi: Neocallimastigomycota. IMA Fungus, 1(2), pp. 181-185. Download the pdf.

Edwards, A, A.M. Anesio, S.M. Rassner, B. Sattler, B. Hubbard, W.T. Perkins, M. Young and G.W. Griffith (2011). Possible interactions between bacterial diversity, microbial activity and supraglacial hydrology of cryoconite holes in Svalbard. ISME Journal (doi:10.1038/ismej.2010.100). Download the pdf.

Griffith, G.W., E. Evans, H.M. Davey & A. Edwards (2011).  Microbiology at the Urdd Eisteddfod.  Microbiology Today, February 2011,  pp. 58-59. Download the pdf.

Griffith, G.W. (2011). Obtaining cultures for teaching praticals in mycology.. Mycologist News, 2011 (1), p5. Download the pdf.

Griffith, G.W. & K. Seifert (2011). Stan Hughes - Order of Canada. Mycologist News, 2011(1), p. 4. Download the pdf.


Griffith, G.W., Baker, S., Fliegerova, K., Liggenstoffer, A, van der Giezen, M., Voigt, K. and Beakes, G. (2010). Anaerobic fungi: Neocallimastigomycota. IMA Fungus, 1(2), pp. 181-185. Download the pdf.

Griffith, G.W. (2010).  On Biodiversity. Letter to New Scientist, 19th May 2010. vol 206 No 2761). p28. Link to the webpage.

Hedger, J.N., J.D. George, G.W. Griffith & D. Peiris (2010).  Lundy fungi: Further studies 2004-2008.  Journal of the Lundy Field Society, 2, pp. 21-44. Download the pdf.

Matheny, P.B. & Griffith, G.W. (2010). Mycoparasitism between Squamanita paradoxa and Cystoderma amianthinum (Cystodermateae, Agaricales). Mycoscience, in press. (10.1007/s10267-010-0052-9). Download the pdf.

Farrell, M., Griffith, G.W., Hobbs, P.J., Perkins, W.T. & Jones, D.L. (2010). Microbial diversity and activity are increased by compost amendment of metal contaminated soil. FEMS MicrobiologyEcology, 71(1), pp. 94-105. Download the pdf.

Farrell, M., Perkins, W.T., Hobbs, P.J., Griffith, G.W. & Jones, D.L. (2010).  Migration of heavy metals in soil as influenced by compost amendments. Environmental Pollution, 158, pp. 55–64. Download the pdf.


Kumar, S., A.K. Puniya, M. Puniya, S.S. Dagar, S.K. Sirohi, K. Singh & G.W. Griffith (2009). Factors affecting rumen methanogens and methane mitigation strategies. World Journal Biotechnology and Microbiology25, pp. 1557-1566. Download the pdf.

Schoch, C.L., G. Sung, F. López-Giráldez, J.P. Townsend, J. Miadlikowska, V. Hofstetter, B. Robbertse, P.B. Matheny, F. Kauff, Z. Wang C. Gueidan, R. M. Andrie, K. Trippe, L.M. Ciufetti, A. Wynns, E. Fraker, B.P. Hodkinson, G. Bonito, R. Yahr, J.Z. Groenewald, M. Arzanlou, G.S. de Hoog, P.W. Crous, D. Hewitt, D.H. Pfister, K. Peterson, M. Gryzenhout, M.J. Wingfield, A. Aptroot, S. Suh, M. Blackwell, D.M. Hillis, G.W. Griffith, L.A. Castlebury, A.Y. Rossman, H. T. Lumbsch, R. Lücking, B. Büdel, A. Rauhut, P. Diederich, D. Ertz, D.M. Geiser, K. Hosaka, P. Inderbitzin, J. Kohlmeyer, B. Volkmann-Kohlmeyer, L. Mostert, K. O'Donnell, H. Sipman, J.D. Rogers, R.A. Shoemaker, J. Sugiyama, R.C. Summerbell, W. Untereiner, P. Johnston, S. Stenroos, A. Zuccaro, P. Dyer, P. Crittenden, M.S. Cole, K. Hansen, J.M. Trappe , F. Lutzoni and J.W. Spatafora (2008). The Ascomycota Tree of Life: A phylum-wide phylogeny clarifies the origin and evolution of fundamental reproductive and ecological traits. Systematic Biology, 58, pp. 224 - 239. Download the pdf.

Griffith, G.W.*, E. Ozkose, M.K. Theodorou and D.R. Davies (2009). Diversity of anaerobic fungal populations in cattle revealed by selective enrichment culture using different carbon sources. Fungal Ecology, 2, pp. 87-97. Download the pdf.


Meinhardt, L.W., J. Rincones, B.A. Bailey, M.C. Aime, G.W. Griffith, D. Zhang and G.A. Pereira. (2008). Moniliophthora perniciosa, the causal agent of witches' broom disease of cacao: What's new from this old foe? Molecular Plant Pathology, 9(5), pp. 577-588. Download the pdf.

Edwards, J.E., A.H. Kingston-Smith, H.R. Jimenez, S.A. Huws, K. Skøt, G.W. Griffith, N.R. McEwan and M.K. Theodorou (2008). Dynamics of initial colonisation of nonconserved perennial ryegrass by anaerobic fungi in the bovine rumen. FEMS Microbiology Ecology, 66 (3), pp. 537-546. Download the pdf.

Delneri, D., D.C. Hoyle, K. Gkargkas, E.J.M. Cross, B. Rash, L. Zeef, H. Leong, H.M. Davey, A. Hayes, D.B. Kell, G.W. Griffith & S.G. Oliver (2008). Identification and characterisation of high flux control genes of Saccharmoyces cerevisiae through competition analysis in continuous cultures. Nature Genetics, 40 (1), pp. 113-117. Download the pdf.

Griffith, G.W. & K. Roderick (2008).  Saprotrophic basidiomycetes in grasslands: distribution and function. In: Ecology of Saprotrophic Basidiomycetes (eds. L. Boddy, J.C. Frankland and P. van Pest).  British Mycological Society Symposia Series. Elsevier Ltd.. pp. 275-297. Download the pdf.

Nagpal, R.  A.K. Puniya, G.W. Griffith, G. Goel, M. Puniya, J.P. Sehgal, K. Singh (2008).  Anaerobic Rumen Fungi: Potential and Applications In: Agriculturally Important Micro-Organisms, Volume 1.  Eds:  G.G. Khachatourians, D.K. Arora, T.P. Rajendran, A.K Srivastava(series editor D.K. Arora).  Academic World International,  Chapter 17, pp.375-393. Download the pdf.


Griffith, G.W., G.L. Easton, A. Detheridge, K. Roderick, A. Edwards, H.J. Worgan, J. Nicholson, & W.T. Perkins. (2007). Copper deficiency in potato dextrose agar causes reduced pigmentation in fungal cultures. FEMS Microbiology Letters. 276 (2), pp. 165-171. Download the pdf.

Hibbett, D. S., Binder, M., Bischoff, J. F., Blackwell, M., Cannon, P. F., Eriksson, O.-E., Huhndorf, S., James, T. Y., Kirk, P. M., Lucking, R., Lumbsch, T., Lutzoni, F., Matheny, P. B., McLaughlin, D. J., Powell, M. J., Redhead, S., Schoch, C. L., Spatafora, J. W., Stalpers, J. A., Vilgalys, R., Aime, M. C., Aptroot, A., Bauer, R., Begerow, D., Benny, G. L., Castlebury, L. A., Crous, P. W., Dai, Y.-C., Gams, W., Geiser, D. M., Griffith, G. W., Gueidan, C., Hawksworth, D. L., Hestmark, G., Hosaka, K., Humber, R. A., Hyde, K., Ironside, J. E., Koljalg, U., Kurtzman, C. P., Larsson, K.-H., Lichwardt, R., Longcore, J., Miadlikowska, J., Miller, A., Moncalvo, J.-M., Mozley-Standridge, S., Oberwinkler, F., Parmasto, E., Reeb, V., Rogers, J. D., Roux, C., Ryvarden, L., Sampaio, J. P., Schussler, A., Sugiyama, J., Thorn, R. G., Tibell, L., Untereiner, W. A., Walker, C., Wang, Z., Weir, A., Weiss, M., White, M. M., Winka, K., Yao, Y.-J. and Zhang, N. (2007). A Higher-Level Phylogenetic Classification of the Fungi. Mycological Research 111, pp. 509-547. Download the pdf.

Griffith, G.W. (2007). Phytophthora: a blight on Ireland. Microbiology Today. 34(1), February 2007. pp.12-15. Download the pdf.

Hedger, J.N., J.D. George, G.W. Griffith & L. Deacon.  (2007).  The Mushrooms and Toadstools of Lundy. Annual Report of the Lundy Field Society, 56, pp. 13-25. Download the pdf (or download the whole issue).


James, T.Y., P.M. Letcher, J.E. Longcore, S.E. Mozley-Standridge, D. Porter, M.J. Powell, G.W. Griffith & R. Vilgalys (2006). Chytridiomycota: 14 clades of flagellated Fungi. Mycologia, 98(6), pp. 860-871. Download the pdf.

James, T.Y., F. Kauff, C. Schoch, P.B. Matheny, V., C.J. Cox, G. Celio, C. Gueidan, E. Fraker, J. Miadlikowska, H.T. Lumbsch, A. Rauhut,V. Reeb, A.E. Arnold, A. Amtoft, J.E. Stajich, K. Hosaka, G. Sung, D. Johnson, B. O'Rourke, M. Crockett, M. Binder, J.M. Curtis, J.C. Slot, Z. Wang, A.W. Wilson, A. Schußler, J.E. Longcore, K. O'Donnell, S. Mozley-Standridge, D. Porter, P.M. Letcher, M.J. Powell, J.W. Taylor, M.M. White, G.W. Griffith, D.R. Davies, R.A. Humber, J.B. Morton, J. Sugiyama, A.Y. Rossman, J.D. Rogers, D.H. Pfister, David Hewitt, K. Hansen, S. Hambleton, R.A. Shoemaker, J. Kohlmeyer, B. Volkmann-Kohlmeyer, R.A. Spotts, M. Serdani, P.W. Crous, K.W. Hughes, K. Matsuura, E. Langer, G. Langer, W.A. Untereiner, R. Lucking, B. Budel, D.M. Geiser, A. Aptroot, P. Diederich, I. Schmitt, M. Schultz, R. Yahr, D.S. Hibbett, F. Lutzoni, D.J. McLaughlin, J.W. Spatafora & R. Vilgalys. (2006). Reconstructing the early evolution of the fungi using a six gene phylogeny. Nature, London (Article), 443, 19th October, pp. 818-822. Download the pdf.

Evans, L., J.N. Hedger, D. Brayford, M. Stavri, E. Smith, G. O'Donnell, A.I. Gray, G.W. Griffith and S. Gibbons (2006). An antibacterial hydroxy fusidic acid analogue from the mitosporic fungus Acremonium crotocinigenum (Schol-Schwarz) W. Gams. Phytochemistry, 67, pp. 2110-2114. Download the pdf.

Rincones, J., G.D. Mazotti, G.W. Griffith, A. Pomela, A. Figueira, G.A. Leal, M.V. Queiroz, J.F. Pereira, R.A. Azevedo, G.A.G. Pereira. & L.W. Meinhardt. (2006). Genetic variability and chromosome-length polymorphisms of the witches' broom pathogen Crinipellis perniciosa from various plant hosts in South America. Mycological Research, 110(7), pp. 821-832. Download the pdf.

Zhao, H., Parry, R.L., Ellis, D.I., Griffith, G.W. & Goodacre, R. (2006). The rapid differentiation of Streptomyces isolates using Fourier transform infrared spectroscopy, Vibrational Spectroscopy, 40, pp. 213-218. Download the pdf.


Johnson, H.E., R.J. Merry, D.B. Kell, M.K. Theodorou & G.W. Griffith (2005). Vacuum packing: A model system for laboratory-scale silage fermentations. Journal of Applied Microbiology, 98(1), pp. 106-113. Download the pdf.

Griffith, G.W. (2005). Waxcap grasslands at Sourhope. In:  The soil beneath our feet.  NERC Soil Biodiversity Programme Final Report. Download the pdf.


Griffith, G.W. (2004). Use of stable isotopes in fungal ecology. Mycologist, 18(4), pp. 177-183. Download the pdf.

Winters, A.L., F.R. Minchin, Z.S. Davies, A.H. Kingston-Smith, M.K. Theodorou, G.W Griffith & R.J. Merry (2004). Effects of manipulating the protein content of white clover on silage quality. Animal Feed Science and Technology, 116(3), pp. 319-331. Download the pdf.

Griffith, G.W. (2004). Witches' brooms and frosty pods: threats to world cacao production. The Biologist, 51(2), pp. 71-75. Download the pdf.

Johnson, H.E., D. Broadhurst, D.B. Kell, M.K. Theodorou, R.J. Merry & G.W. Griffith (2004). High-throughput metabolic fingerprinting of legume silage fermentations using Fourier transform infrared spectroscopy and chemometrics. Applied and Environmental Microbiology, 70(3), pp.1583-1592. Download the pdf.

Griffith, G.W., Bratton, J.L. & Easton, G. 2004. Charismatic megafungi: the conservation of waxcap grasslands. British Wildlife, 15(3), pp. 31-43. Download the pdf.

Griffith, G.W. (2004). Microbiology in the Regions. Report of the 2nd All-Wales Microbiology Meeting, Gregynog Hall, 15-17 March 2004. Microbiology Today, 31(4) November,  p. 191. Download the pdf.

Griffith, G.W. (2004). Ymchwil cacao.  Cysylltiadau rhwng Aber a Brasil.  PROM, 2004, pp.18-20. Download the pdf.

Griffith, G.W. (2004).  A mycologist in Westminster.  Microbiology Today, 31(1) February,  pp. 28-29. Download the pdf.


Scullion, J., G.N. Elliott, W.E. Huang, R. Goodacre, H.J. Worgan, R. Darby, M.J. Bailey, D. Gwynn-Jones, G.W. Griffith, M.K. Winson, P.A. Williams, C. Clegg & J. Draper (2003) Use of earthworm casts to validate FTIR spectroscopy as a 'sentinel' technology for high-throughput monitoring of global changes in microbial ecology. Pedobiologia. 47, pp. 440-446. Download the pdf.

Scullion, J., R. Goodacre, G.N. Elliott, W.E. Huang, H.J. Worgan, D. Gwynn-Jones, G.W. Griffith, R. Darby, M.J. Bailey, C. Clegg & J. Draper (2003). Food quality and microbial succession in ageing earthworm casts: standard microbial indices and metabolic fingerprinting. Pedobiologia. 47, pp. 888-894. Download the pdf.

Griffith, G.W., J.N. Nicholson, A. Nenninger, R.N. Birch & J.N. Hedger (2003). Witches' brooms and frosty pods: two major pathogens of cacao. New Zealand Journal of Botany, 41, pp. 423-435. Download the pdf.

Wilberforce, E.M, L. Boddy, R. Griffiths & G.W. Griffith (2003). Agricultural management affects diversity of culturable root endophytes from mesotrophic grasslands in Wales. Soil Biology and Biochemistry. 35, pp. 1143-1154. Download the pdf.

Griffith, G.W. & G.R. Easton (2003). Ydi'r balerina binc yn dawnsio yn eich gardd chi?  Y Naturiaethwr, 2(13), pp. 48-49. Download the pdf.


Griffith, G.W., G.L. Easton & A.W. Jones (2002). Ecology and Diversity of Waxcap (Hygrocybe spp.) Fungi. Botanical Journal of Scotland, 54(1), pp.7-22. Download the pdf.

Griffith, G.W., J.P. Day & D.B. Kell (2002). Use of flow cytometry in the detection of plant pathogenic spores. Proceedings of the British Crop Protection Council conference, Brighton, 18-21st November, pp. 417-424. Download the pdf.

Day, J.P., Kell, D.B. and Griffith, G.W. (2002). Differentiation of Phytophthora infestans sporangia from other airborne biological particles using flow cytometry. Applied and Environmental Microbiology. 68(1), pp. 37-45. Download the pdf.

Thomas, M., Keirle, K., Griffith G.W., Hughes S.G., Hart, P.M. & Schollar, J. (2002). The Biotechnology Summer School: a novel teaching initiative.Innovations in Education and Teaching International, 39(2), pp. 124-136. Download the pdf.


Ozkose, E., Thomas, B.J., Davies, D.R., Griffith, G.W. and Theodorou, M.K. (2001). Cyllamyces aberensis gen. nov., sp. nov., an anaerobic gut fungus from cattle with bulbous holdfast, branched sporangiophores and polycentric development. Canadian Journal of Botany, 79, pp. 666-673. Download the pdf.

M. Thomas, S.G. Hughes, P.M. Hart, J. Schollar, K. Keirle and G. W. Griffith. (2001). Group project work in biotechnology and its impact on key skills. Journal of Biological Education, 34 (3). pp.133-140. Download the pdf.

Eason, W.R., K.J. Webb, T.P.T. Michaelson-Yeates, M.T. Abberton, G.W. Griffith, C.M. Culshaw, J.E. Hooker and M.S. Dhanoa (2001). Effect of genotype of Trifolium repens on mycorrhizal symbiosis with Glomus mosseae. Journal of Agricultural Science, Cambridge, 137(1), pp. 27-36. Download the pdf.

Griffith, G.W.
(2001).  Spore counts could predict blight spread. Potato ReviewSeptember 2001, pp. 4-5.
Download the pdf.


Griffith, G.W.. (2000). Application of molecular biology to cocoa pathology. Cocoa Growers’ Bulletin, 52, pp. 46-57. Download the pdf

Davies, Z.S., D. Mason, A.E. Brooks, G.W. Griffith, R.J. Merry & M.K. Theodorou (2000).  An automated system for measuring gas production from forages inoculated with rumen fluid and its use in determining the effect of enzymes on grass silage. Animal Feed Science and Technology.  84. pp. 1-16.  Download the pdf

Davies, Z.S., Gilbert, R.C., Merry R.J., Kell, D.B., Theodorou, M.K. & Griffith, G.W. (2000).  Efficient improvement of silage additives using genetic algorithms. Applied and Environmental Microbiology.  66(4),  1435-1443. Download the pdf


Griffith, G.W., M.S. Stark and A.J. Clutterbuck (1999).  Wild type and mutant alleles of Aspergillus nidulans developmental regulator gene brlA: correlation of variant sites with protein function.  Molecular and General Genetics.  262. pp. 892-897. Download the pdf.

McGranaghan, P., J.C. Davies,, G.W. Griffith , D.R. Davies & M.K. Theodorou (1999).  The survival of anaerobic fungi in cattle faeces. FEMS Microbiology Ecology. 29(3),  pp. 293-300. Download the pdf.

Griffith, G.W & D.S. Shaw (1998). Polymorphisms in Phytophthora infestans: Four mitochondrial haplotypes are detected after PCR amplification of DNA from pure cultures or from host lesions. Applied and Environmental Microbiology, 64 (10) pp. 4007-4014. Download the pdf.

Griffith, G.W. & J.P. Day (1995).  Herio'r malltod tatws.  Delta.  3,  pp.14-16 [In Welsh].

Griffith, G.W., R. Snell & D.S. Shaw (1995).  Late blight (Phytophthora infestans) on tomato in the tropics. The Mycologist. 9 (2),  pp. 87-89. Download the pdf.

Griffith, G.W., G.I. Jenkins, E.J. Milner-White & A.J. Clutterbuck (1994).  Similarity at the amino acid level between plant phytochromes and a regulator of asexual sporulation in Emericella (=Aspergillus) nidulans.  Photochemistry and Photobiology,  59(2),  pp. 252-256. Download the pdf.

Griffith, G.W. & J.N. Hedger (1994).  The breeding biology of biotypes of the witches' broom pathogen of cocoa, Crinipellis perniciosa.  Heredity, 72,  pp. 278-289. Download the pdf.

Griffith, G.W., E. Bravo, F.J. Wilson, D.M. Lewis & J.N. Hedger (1994).  Autecology and evolution of the witches' broom pathogen (Crinipellis perniciosa.) of cocoa.  Chapter 15 in Ecology of Plant Pathogens.(Eds. J.P. Blakeman & B. Williamson), CAB International, Wallingford.  pp. 245-267.

Griffith, G.W. (1994).  Phenoloxidases.  Chapter 28 in Aspergillus nidulans: 50 years on (Eds S.D. Martinelli and J.R. Kinghorn); Progress in Industrial Microbiology, Volume 29.  Elsevier Science Publishers, Amsterdam.  pp. 763-788. Download the pdf

Griffith, G.W. & J.N. Hedger (1994).  Spatial distribution of mycelia of the liana biotype of Crinipellis perniciosa Stahel (Sing.) in tropical forest.  New Phytologist 127 (2).  pp. 243-259. Download the pdf.

Griffith, G.W., R. Snell, J.R. Witcombe, D.S. Shaw, L.L. Black & Y-H. Huang (1994).  Host-pathogen genetics of late-blight in tomato.  Compendium of Current Research in Plant Sciences.  (Eds. J.R. Witcombe & D. Wright).  Overseas Development Administration.  pp. 67-69.

Davies, R.W., A.B. Roberts, A.J. Morris, G.W. Griffith, J. Jerecic, S. Gandhi, K. Kaiser & A. Savioz (1994).  Enhanced access to rare brain cDNAs by prescreening libraries:  206 new mouse brain ESTs.  Genomics.  24 (3).  pp. 456-463. Download the pdf.

Griffith, G.W. & J.N. Hedger (1994).  Dual culture of Crinipellis perniciosa and potato callus.  European Journal of Plant Pathology.  100 (6).  pp. 371-379. Download the pdf.

Griffith, G.W., F.J. Wilson & J.N. Hedger (1994).  Population structure, breeding biology and evolutionary relationships in the biotypes of the witches' broom disease fungus, Crinipellis perniciosa, in South America  pp.45-52.  Proceedings of the 11th International Cocoa Research Conference, Yamoussoukro, Ivory Coast, 18-24th July, 1993.  Cocoa Producers' Alliance.  Lagos, Nigeria.  

Griffith, G.W. & J.N. Hedger (1993).  A novel method for fruiting the cocoa pathogen Crinipellis perniciosa using a bran-vermiculite medium.  Netherlands Journal of Plant Pathology,  99,  pp. 227-230. Download the pdf

Griffith, G.W. (1988).  Clefyd ysgubell y wrach.  Y Gwyddonydd, 25, pp. 105-107. [In Welsh]

Hedger, J.N., G.W. Griffith, H.E. Brownlee, A.R. McEuen, E. Bravo & I.M. Scott (1988).  Witches' brooms and cocoa.  Journal of Agriculture of the University College of Wales., 51, pp. 164-183.

Lowe, S.E., G.W. Griffith, A. Milne, M.K. Theodorou & A.P.J. Trinci (1987).  The life cycle and growth kinetics of an anaerobic rumen fungus.  Journal of General Microbiology, 133, 1815-1827. Download the pdf.


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Aberystwyth University HomepageAberystwyth Bio Centre   Institute of Biological Sciences
Last Updated February 3, 2012 Designed and Developed by Sharon Nicholson


This Page Is Currently Under Construction




Top of Page

Aberystwyth University HomepageAberystwyth Bio Centre   Institute of Biological Sciences
Last Updated February 3, 2012 Designed and Developed by Sharon Nicholson
Designed and Developed by Sharon Nicholson